D. Bakshi, P. Singhal, S. K. Mahajan, P. Subramaniam, U. Tuteja et al., Development of a real-time PCR assay for the diagnosis of scrub typhus cases in India and evidence of the prevalence of new genotype of O. tsutsugamushi, Acta Tropica, vol.104, issue.1, pp.63-71, 2007.
DOI : 10.1016/j.actatropica.2007.07.013

S. D. Blacksell, R. Luksameetanasan, T. Kalambaheti, N. Aukkanit, D. H. Paris et al., isolates causing human scrub typhus at two sites in north-eastern and western Thailand, FEMS Immunology & Medical Microbiology, vol.52, issue.3, pp.335-342, 2008.
DOI : 10.1111/j.1574-695X.2007.00375.x

F. M. Bozeman and B. L. Elisberg, Serological Diagnosis of Scrub Typhus by Indirect Immunofluorescence, Experimental Biology and Medicine, vol.112, issue.3, pp.568-573, 1963.
DOI : 10.3181/00379727-112-28107

M. Cao, H. Guo, T. Tang, C. Wang, X. Li et al., Spring Scrub Typhus, People's Republic of China, Emerging Infectious Diseases, vol.12, issue.7, pp.1463-1465, 2006.
DOI : 10.3201/eid1209.060257

S. Chattopadhyay and A. L. Richards, Scrub Typhus Vaccines: Past History and Recent Developments, Human Vaccines, vol.3, issue.3, pp.73-80, 2007.
DOI : 10.4161/hv.3.3.4009

N. H. Cho, H. R. Kim, J. H. Lee, S. Y. Kim, J. Kim et al., The Orientia tsutsugamushi genome reveals massive proliferation of conjugative type IV secretion system and host cell interaction genes, Proc. Natl. Acad. Sci. U.S.A. 104, pp.7981-7986, 2007.
DOI : 10.1073/pnas.0611553104

R. E. Coleman, T. Monkanna, K. J. Linthicum, D. A. Strickman, S. P. Frances et al., Occurrence of Orientia tsutsugamushi in small mammals from Thailand, Am. J. Trop. Med. Hyg, vol.69, pp.519-524, 2003.

A. L. Corwin, W. Soeprapto, P. S. Widodo, E. Rahardjo, D. J. Kelly et al., Short report: surveillance of rickettsial infections in Indonesian military personnel during peace keeping operations in Cambodia, Am. J. Trop. Med. Hyg, vol.57, pp.569-570, 1997.

D. Jr, J. J. Russell, and P. K. , An analysis of fevers of unknown origin in American soldiers in Vietnam, Ann. Intern. Med, vol.66, pp.1129-1143, 1967.

V. Duong, K. Blassdell, T. T. May, L. Sreyrath, L. Gavotte et al., Diversity of Orientia tsutsugamushi clinical isolates in Cambodia reveals active selection and recombination process, Infection, Genetics and Evolution, vol.15, 2010.
DOI : 10.1016/j.meegid.2010.08.015
URL : https://hal.archives-ouvertes.fr/pasteur-00808212

R. C. Edgar, MUSCLE: a multiple sequence alignment method with reduced time and space complexity, BMC Bioinformatics, vol.5, issue.1, p.113, 2004.
DOI : 10.1186/1471-2105-5-113

B. L. Elisberg, V. Sangkasuvana, J. M. Campbell, F. M. Bozeman, and P. Bodhidatta, Physiogeographic distribution of scrub typhus in Thailand, Acta Med. Biol, vol.15, pp.61-67, 1967.

T. Enatsu, H. Urakami, and A. Tamura, strains based on the sequence homologies of 56-kDa type-specific antigen genes, FEMS Microbiology Letters, vol.180, issue.2, pp.163-169, 1999.
DOI : 10.1111/j.1574-6968.1999.tb08791.x

J. Felsenstein, Confidence Limits on Phylogenies: An Approach Using the Bootstrap, Evolution, vol.39, issue.4, pp.783-791, 1985.
DOI : 10.2307/2408678

Y. Furuya, Y. Yoshida, T. Katayama, F. Kawamori, S. Yamamoto et al., Specific amplification of Rickettsia tsutsugamushi DNA from clinical specimens by polymerase chain reaction, J. Clin. Microbiol, vol.29, pp.2628-2630, 1991.

N. Galtier, M. Gouy, and C. Gautier, SEAVIEW and PHYLO_WIN: two graphic tools for sequence alignment and molecular phylogeny, Bioinformatics, vol.12, issue.6, pp.543-548, 1996.
DOI : 10.1093/bioinformatics/12.6.543
URL : https://hal.archives-ouvertes.fr/hal-00435028

M. Gouy, S. Guindon, and O. Gascuel, SeaView Version 4: A Multiplatform Graphical User Interface for Sequence Alignment and Phylogenetic Tree Building, Molecular Biology and Evolution, vol.27, issue.2, pp.221-224, 2010.
DOI : 10.1093/molbev/msp259
URL : https://hal.archives-ouvertes.fr/lirmm-00705187

T. A. Hall, BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95, NT. Nucl. Acids Symp. Ser, vol.98, issue.41, pp.95-98, 1999.

B. Hanson, Identification and partial characterization of Rickettsia tsutsugamushi major protein immunogens, Infect. Immun, vol.50, pp.603-609, 1985.

M. Jensenius, P. E. Fournier, and D. Raoult, Rickettsioses and the international traveler, Clin. Infect. Dis, vol.39, pp.1493-1499, 2004.

T. Katayama, M. Hara, Y. Furuya, T. Nikkawa, and H. Ogasawara, Scrub typhus (tsutsugamushi disease) in Kanagawa Prefecture, Jpn. J. Infect. Dis, vol.59, pp.207-208, 2001.

B. Khuntirat, K. Lerdthusnee, W. Leepitakrat, A. Kengluecha, K. Wongkalasin et al., Isolated from Wild-Caught Rodents and Chiggers in Northern Thailand, Annals of the New York Academy of Sciences, vol.39, issue.1, pp.205-212, 2003.
DOI : 10.1111/j.1749-6632.2003.tb07364.x

J. H. Lee, N. H. Cho, S. Y. Kim, S. Y. Bang, H. Chu et al., into Host Cells through Interaction with a 56???kDa Type???Specific Antigen, The Journal of Infectious Diseases, vol.198, issue.2, pp.250-257, 2008.
DOI : 10.1086/589284

K. Nakayama, A. Yamashita, K. Kurokawa, T. Morimoto, M. Ogawa et al., The Whole-genome Sequencing of the Obligate Intracellular Bacterium Orientia tsutsugamushi Revealed Massive Gene Amplification During Reductive Genome Evolution, DNA Research, vol.15, issue.4, pp.185-199, 2008.
DOI : 10.1093/dnares/dsn011

M. Nei and S. Kumar, Molecular Evolution and Phylogenetics, 2000.

N. Ohashi, H. Nashimoto, H. Ikeda, and A. Tamura, Cloning and sequencing of the gene (tsg56) encoding a type-specific antigen from Rickettsia tsutsugamushi, Gene, vol.91, issue.1, pp.119-122, 1990.
DOI : 10.1016/0378-1119(90)90171-M

N. Ohashi, H. Nashimoto, H. Ikeda, and A. Tamura, Diversity of immunodominant 56-kDa type-specific antigen (TSA) of Rickettsia tsutsugamushi, Sequence and comparative analyses of the genes encoding TSA homologues from four antigenic variants, J. Biol. Chem, vol.267, pp.12728-12735, 1992.

N. Ohashi, A. Tamura, M. Ohta, and K. Hayashi, Purification and partial characterization of a type-specific antigen of Rickettsia tsutsugamushi, Infect. Immun, vol.57, pp.1427-1431, 1989.

D. H. Paris, N. Aukkanit, K. Jenjaroen, S. D. Blacksell, and N. P. Day, A highly sensitive quantitative real-time PCR assay based on the groEL gene of contemporary Thai strains of Orientia tsutsugamushi, Clinical Microbiology and Infection, vol.15, issue.5, pp.488-495, 2009.
DOI : 10.1111/j.1469-0691.2008.02671.x

P. Parola, S. D. Blacksell, R. Phetsouvanh, S. Phongmany, J. M. Rolain et al., from Humans with Scrub Typhus, Laos, Emerging Infectious Diseases, vol.14, issue.9, pp.1483-1485, 2008.
DOI : 10.3201/eid1409.071259

S. R. Paul, S. Karanth, and C. Dickson, Scrub Typhus along the Thai-Kampuchean Border: New Treatment Regimen, Tropical Doctor, vol.66, issue.3, pp.104-107, 1987.
DOI : 10.1177/004947558701700304

Y. Qiang, A. Tamura, H. Urakami, Y. Makisaka, S. Koyama et al., Isolated in Taiwan According to the Sequence Homologies of 56-kDa Type-Specific Antigen Genes, Microbiology and Immunology, vol.51, issue.8, pp.577-583, 2003.
DOI : 10.1111/j.1348-0421.2003.tb03420.x

H. I. Ree, T. E. Kim, I. Y. Lee, S. H. Jeon, U. W. Hwang et al., Determination and geographical distribution of Orientia tsutsugamushi serotypes in Korea by nested polymerase chain reaction, Am. J. Trop. Med. Hyg, vol.65, pp.528-534, 2001.

N. Saitou and M. Nei, The neighbor-joining method: a new method for reconstructing phylogenetic trees, Mol. Biol. Evol, vol.4, pp.406-425, 1987.

S. Y. Seong, M. S. Huh, W. J. Jang, S. G. Park, J. G. Kim et al., Induction of homologous immune response to Rickettsia tsutsugamushi Boryong with a partial 56-kilodalton recombinant antigen fused with the maltose-binding protein MBP-Bor56, Infect. Immun, vol.65, pp.1541-1545, 1997.

S. Y. Seong, H. R. Kim, M. S. Huh, S. G. Park, J. S. Kang et al., Induction of neutralizing antibody in mice by immunization with recombinant 56 kDa protein of Orientia tsutsugamushi, Vaccine, vol.15, issue.16, pp.1741-1747, 1997.
DOI : 10.1016/S0264-410X(97)00112-6

S. Y. Seong, M. K. Kim, S. M. Lee, Z. Odgerel, M. S. Choi et al., Neutralization epitopes on the antigenic domain II of the Orientia tsutsugamushi 56-kDa protein revealed by monoclonal antibodies, Vaccine, vol.19, issue.1, pp.2-9, 2000.
DOI : 10.1016/S0264-410X(00)00167-5

A. Shirai, D. M. Robinson, G. W. Brown, E. Gan, and D. L. Huxsoll, ANTIGENIC ANALYSIS BY DIRECT IMMUNOFLUORESCENCE OF 114 ISOLATES OF RICKETTSIA TSUTSUGAMUSHI RECOVERED FROM FEBRILE PATIENTS IN RURAL MALAYSIA, Japanese Journal of Medical Science and Biology, vol.32, issue.6, pp.337-344, 1979.
DOI : 10.7883/yoken1952.32.337

A. Shirai, W. Jr, and C. L. , Serologic classification of scrub typhus isolates from Pakistan, Am. J. Trop. Med. Hyg, vol.24, pp.145-153, 1975.

A. Shishido, STRAIN VARIATION OF RICKETTSIA ORIENTALIS IN THE COMPLEMENT FIXATION TEST*, Japanese Journal of Medical Science and Biology, vol.17, issue.2-3, pp.59-72, 1964.
DOI : 10.7883/yoken1952.17.59

P. Sonthayanon, S. J. Peacock, W. Chierakul, V. Wuthiekanun, S. D. Blacksell et al., High Rates of Homologous Recombination in the Mite Endosymbiont and Opportunistic Human Pathogen Orientia tsutsugamushi, PLoS Neglected Tropical Diseases, vol.31, issue.7, p.752, 2010.
DOI : 10.1371/journal.pntd.0000752.s001

C. K. Stover, D. P. Marana, J. M. Carter, B. A. Roe, E. Mardis et al., The 56- kilodalton major protein antigen of Rickettsia tsutsugamushi: molecular cloning and sequence analysis of the sta56 gene and precise identification of a strainspecific epitope, Infect. Immun, vol.58, pp.2076-2084, 1990.

C. K. Stover, D. P. Marana, G. A. Dasch, and E. V. Oaks, Molecular cloning and sequence analysis of the Sta58 major antigen gene of Rickettsia tsutsugamushi: sequence homology and antigenic comparison of Sta58 to the 60-kilodalton family of stress proteins, Infect. Immun, vol.58, pp.1360-1368, 1990.

A. Tamura, N. Ohashi, H. Urakami, and S. Miyamura, Classification of Rickettsia tsutsugamushi in a New Genus, Orientia gen. nov., as Orientia tsutsugamushi comb. nov., International Journal of Systematic Bacteriology, vol.45, issue.3, pp.589-591, 1995.
DOI : 10.1099/00207713-45-3-589

A. Tamura, N. Ohashi, H. Urakami, K. Takahashi, and M. Oyanagi, Analysis of polypeptide composition and antigenic components of Rickettsia tsutsugamushi by polyacrylamide gel electrophoresis and immunoblotting, Infect. Immun, vol.48, pp.671-675, 1985.

S. T. Tay, Y. M. Rohani, T. M. Ho, and D. Shamala, Strains in Malaysia, Microbiology and Immunology, vol.71, issue.1, pp.67-71, 2005.
DOI : 10.1111/j.1348-0421.2005.tb03641.x

M. M. Thiebaut, F. Bricaire, and D. Raoult, Scrub Typhus after a Trip to Vietnam, New England Journal of Medicine, vol.336, issue.22, pp.1613-1614, 1997.
DOI : 10.1056/NEJM199705293362220

R. Traub, W. Jr, and C. L. , CURRENT CONCEPTS OF THE ECOLOGY OF CHIGGER-BORNE RICKETTSIOSIS (SCRUB TYPHUS)*, Japanese Journal of Medical Science and Biology, vol.27, issue.1, pp.1-5, 1974.
DOI : 10.7883/yoken1952.27.1

H. Wilde, J. Pornsilapatip, T. Sokly, and S. Thee, Murine and scrub typhus at Thai- Kampuchean border displaced persons camps, Trop. Geogr. Med, vol.43, pp.363-369, 1991.