J. Schimel, In Ecological Studies: Analysis and Synthesis, pp.239-254, 1995.

A. Jumpponen, Soil fungal community assembly in a primary successional glacier forefront ecosystem as inferred from rDNA sequence analyses, New Phytologist, vol.21, issue.3, pp.569-578, 2003.
DOI : 10.1046/j.1469-8137.2003.00767.x

A. Zumsteg, E. Bååth, B. Stierli, J. Zeyer, and B. Frey, Bacterial and fungal community responses to reciprocal soil transfer along a temperature and soil moisture gradient in a glacier forefield, Soil Biology and Biochemistry, vol.61, pp.121-132, 2013.
DOI : 10.1016/j.soilbio.2013.02.017

J. A. Robbins and J. A. Matthews, Regional Variation in Successional Trajectories and Rates of Vegetation Change on Glacier Forelands in South-Central Norway, Arctic, Antarctic, and Alpine Research, vol.42, issue.3, pp.351-361, 2010.
DOI : 10.1657/1938-4246-42.3.351

J. Svoboda and G. Henry, Succession in Marginal Arctic Environments, Arctic and Alpine Research, vol.19, issue.4, pp.373-384, 1987.
DOI : 10.2307/1551402

M. Davey, (L.) Delabre (Polygonaceae) root-associated fungi mirrors plant succession in two glacial chronosequences, Environmental Microbiology, vol.63, issue.8, pp.2777-2790, 2015.
DOI : 10.1111/1462-2920.12770

URL : https://hal.archives-ouvertes.fr/hal-01205036

A. E. Vater and J. A. Matthews, Testing the 'addition and persistence model' of invertebrate succession in a subalpine glacier-foreland chronosequence: Fabergstolsbreen, southern Norway, The Holocene, vol.23, issue.8, pp.1151-1162, 2013.
DOI : 10.1177/0959683613483623

L. R. Walker and R. Del-moral, Primary succession and ecosystem rehabilitation, 2003.
DOI : 10.1017/CBO9780511615078

R. Blaalid, Changes in the root-associated fungal communities along a primary succession gradient analysed by 454 pyrosequencing, Molecular Ecology, vol.187, issue.8, pp.1897-1908, 2012.
DOI : 10.1111/j.1365-294X.2011.05214.x

S. P. Brown and A. Jumpponen, Contrasting primary successional trajectories of fungi and bacteria in retreating glacier soils, Molecular Ecology, vol.63, issue.2, pp.481-497, 2014.
DOI : 10.1111/mec.12487

S. P. Brown and A. Jumpponen, Phylogenetic diversity analyses reveal disparity between fungal and bacterial communities during microbial primary succession, Soil Biology and Biochemistry, vol.89, pp.52-60, 2015.
DOI : 10.1016/j.soilbio.2015.06.025

O. Mühlmann and U. Peintner, Ectomycorrhiza of Kobresia myosuroides at a primary successional glacier forefront, Mycorrhiza, vol.16, issue.5, pp.355-362, 2008.
DOI : 10.1007/s00572-008-0188-z

S. Mundra, do not respond to the same fine-scale edaphic gradients as the aboveground vegetation, New Phytologist, vol.37, issue.4, pp.1587-1597, 2015.
DOI : 10.1111/nph.13216

C. L. Lauber, M. S. Strickland, M. A. Bradford, and N. Fierer, The influence of soil properties on the structure of bacterial and fungal communities across land-use types, Soil Biology and Biochemistry, vol.40, issue.9, pp.2407-2415, 2008.
DOI : 10.1016/j.soilbio.2008.05.021

U. N. Nielsen, G. H. Osler, C. D. Campbell, D. F. Burslem, and R. Van-der-wal, The influence of vegetation type, soil properties and precipitation on the composition of soil mite and microbial communities at the landscape scale, Journal of Biogeography, vol.11, issue.7, pp.1317-1328, 2010.
DOI : 10.1111/j.1365-2699.2010.02281.x

A. Smith, E. Marín-spiotta, M. De-graaff, and T. Balser, Microbial community structure varies across soil organic matter aggregate pools during tropical land cover change, Soil Biology and Biochemistry, vol.77, pp.292-303, 2014.
DOI : 10.1016/j.soilbio.2014.05.030

S. Yarwood, A. Wick, M. Williams, and W. L. Daniels, Parent Material and Vegetation Influence Soil Microbial Community Structure Following 30-Years of Rock Weathering and Pedogenesis, Microbial Ecology, vol.12, issue.2, pp.383-394, 2015.
DOI : 10.1007/s00248-014-0523-1

A. Zumsteg, Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier, Microbial Ecology, vol.66, issue.3, pp.552-564, 2012.
DOI : 10.1007/s00248-011-9991-8

P. Kennedy, Ectomycorrhizal fungi and interspecific competition: species interactions, community structure, coexistence mechanisms, and future research directions, New Phytologist, vol.9, issue.4, pp.895-910, 2010.
DOI : 10.1111/j.1469-8137.2010.03399.x

C. W. Welden and W. L. Slauson, The Intensity of Competition Versus its Importance: An Overlooked Distinction and Some Implications, The Quarterly Review of Biology, vol.61, issue.1, pp.23-44, 1986.
DOI : 10.1086/414724

M. N. Thormann, R. S. Currah, and S. E. Bayley, Succession of microfungal assemblages in decomposing peatland plants, Plant and Soil, vol.250, issue.2, pp.323-333, 2003.
DOI : 10.1023/A:1022845604385

K. Nara, H. Nakaya, B. Wu, Z. Zhou, and T. Hogetsu, Underground primary succession of ectomycorrhizal fungi in a volcanic desert on Mount Fuji, New Phytologist, vol.154, issue.3, pp.743-756, 2003.
DOI : 10.1046/j.1469-8137.2003.00844.x

B. Wu, T. Hogetsu, K. Isobe, and R. Ishii, Community structure of arbuscular mycorrhizal fungi in a primary successional volcanic desert on the southeast slope of Mount Fuji, Mycorrhiza, vol.12, issue.6, pp.495-506, 2007.
DOI : 10.1007/s00572-007-0114-9

J. Pohjanen, Interaction with ectomycorrhizal fungi and endophytic Methylobacterium affects nutrient uptake and growth of pine seedlings in vitro, Tree Physiology, vol.34, issue.9, pp.993-1005, 2014.
DOI : 10.1093/treephys/tpu062

A. R. Jumpponen and J. M. Trappe, Dark septate endophytes: a review of facultative biotrophic root-colonizing fungi, New Phytologist, vol.140, issue.2, pp.295-310, 1998.
DOI : 10.1046/j.1469-8137.1998.00265.x

J. Miadlikowska, New insights into classification and evolution of the Lecanoromycetes (Pezizomycotina, Ascomycota) from phylogenetic analyses of three ribosomal RNA- and two protein-coding genes, Mycologia, vol.98, issue.6, pp.1088-1103, 2006.
DOI : 10.1080/15572536.2006.11832636

L. Selbmann, Biodiversity, evolution and adaptation of fungi in extreme environments, Plant Biosystems - An International Journal Dealing with all Aspects of Plant Biology, vol.5, issue.2, pp.237-246, 2013.
DOI : 10.1007/s00248-010-9651-4

J. Rousk, Soil bacterial and fungal communities across a pH gradient in an arable soil, The ISME Journal, vol.61, issue.10, pp.1340-1351, 2010.
DOI : 10.1016/0168-1605(91)90063-U

M. Miransari, Arbuscular mycorrhizal fungi and nitrogen uptake, Archives of Microbiology, vol.108, issue.2, pp.77-81, 2011.
DOI : 10.1007/s00203-010-0657-6

L. He and Y. Tang, Soil development along primary succession sequences on moraines of Hailuogou Glacier, Gongga Mountain, vol.72, pp.259-269, 2008.

C. Kabala and J. Zapart, Initial soil development and carbon accumulation on moraines of the rapidly retreating Werenskiold Glacier, SW Spitsbergen, Svalbard archipelago, Geoderma, vol.175, issue.176, pp.9-20, 2012.
DOI : 10.1016/j.geoderma.2012.01.025

M. Fujiyoshi, Successional changes in ectomycorrhizal fungi associated with the polar willow Salix polaris in a deglaciated area in the High Arctic, Svalbard, Polar Biology, vol.1, issue.5, pp.667-673, 2011.
DOI : 10.1007/s00300-010-0922-9

D. Tilman, Praktische Zahnmedizin Odonto-Stomatologie Pratique Practical Dental Medicine, pp.327-344, 1994.

G. A. Jones and G. H. Henry, Primary plant succession on recently deglaciated terrain in the Canadian High Arctic, Journal of Biogeography, vol.6, issue.2, pp.277-296, 2003.
DOI : 10.1046/j.1365-2699.2003.00818.x

J. Grime, Primary strategies in plants, Transactions of the Botanical Society of Edinburgh, vol.250, issue.2, pp.151-160, 1979.
DOI : 10.2307/1942191

R. B. Mckane, Resource-based niches provide a basis for plant species diversity and dominance in arctic tundra, Nature, vol.415, issue.6867, pp.68-71, 2002.
DOI : 10.1038/415068a

D. L. Taylor, A first comprehensive census of fungi in soil reveals both hyperdiversity and fine-scale niche partitioning, Ecological Monographs, vol.84, issue.1, pp.3-20, 2014.
DOI : 10.2307/1934875

M. Moreau, D. Laffly, D. Joly, and T. Brossard, Analysis of plant colonization on an arctic moraine since the end of the Little Ice Age using remotely sensed data and a Bayesian approach, Remote Sensing of Environment, vol.99, issue.3, pp.244-253, 2005.
DOI : 10.1016/j.rse.2005.03.017

URL : https://hal.archives-ouvertes.fr/halshs-00468659

P. D. Schloss, Introducing mothur: Open-Source, Platform-Independent, Community-Supported Software for Describing and Comparing Microbial Communities, Applied and Environmental Microbiology, vol.75, issue.23, pp.7537-7541, 2009.
DOI : 10.1128/AEM.01541-09

R. C. Edgar, B. J. Haas, J. C. Clemente, C. Quince, and R. Knight, UCHIME improves sensitivity and speed of chimera detection, Bioinformatics, vol.27, issue.16, pp.2194-2200, 2011.
DOI : 10.1093/bioinformatics/btr381

S. P. Brown, Scraping the bottom of the barrel: are rare high throughput sequences artifacts? Fungal Ecol, pp.221-225, 2015.

R. C. Edgar, Search and clustering orders of magnitude faster than BLAST, Bioinformatics, vol.26, issue.19, pp.2460-2461, 2010.
DOI : 10.1093/bioinformatics/btq461

K. Abarenkov, The UNITE database for molecular identification of fungi - recent updates and future perspectives, New Phytologist, vol.51, issue.2, pp.281-285, 2010.
DOI : 10.1111/j.1469-8137.2009.03160.x

L. Tedersoo, T. W. May, and M. E. Smith, Ectomycorrhizal lifestyle in fungi: global diversity, distribution, and evolution of phylogenetic lineages, Mycorrhiza, vol.92, issue.8, pp.217-263, 2010.
DOI : 10.1007/s00572-009-0274-x

F. Meyer, The metagenomics RAST server ??? a public resource for the automatic phylogenetic and functional analysis of metagenomes, BMC Bioinformatics, vol.9, issue.1, p.386, 2008.
DOI : 10.1186/1471-2105-9-386

D. Roberts and . Labdsv, Ordination and multivariate analysis for ecology. R package version 1 Available at: https://cran.r-project, p.17, 2007.

R. Logares, Biogeography of bacterial communities exposed to progressive long-term environmental change, The ISME Journal, vol.4, issue.5, pp.937-948, 2013.
DOI : 10.1016/S0012-821X(97)00204-5

M. A. Rodríguez-gironés and L. Santamaría, A new algorithm to calculate the nestedness temperature of presence-absence matrices, Journal of Biogeography, vol.90, issue.5, pp.924-935, 2006.
DOI : 10.1007/s004420050348

M. Geel, A. Ceustermans, W. Hemelrijck, B. Lievens, and O. Honnay, Decrease in diversity and changes in community composition of arbuscular mycorrhizal fungi in roots of apple trees with increasing orchard management intensity across a regional scale, Molecular Ecology, vol.25, issue.4, pp.941-952, 2015.
DOI : 10.1111/mec.13079

J. Mcdonald, 26360 | DOI: 10.1038/srep26360 53, Handbook of biological statistics 2nd edn, pp.221-224, 2009.

P. ?milauer and J. Lep?, Multivariate analysis of ecological data using CANOCO 5, 2014.
DOI : 10.1017/CBO9781139627061

T. Braak and C. J. , Canonical Correspondence Analysis: A New Eigenvector Technique for Multivariate Direct Gradient Analysis, Ecology, vol.67, issue.5, pp.1167-1179, 1986.
DOI : 10.2307/1938672