A. Renvoisé and D. Raoult, L???actualit?? des rickettsioses, M??decine et Maladies Infectieuses, vol.39, issue.2, pp.71-81, 2009.
DOI : 10.1016/j.medmal.2008.11.003

C. Keller, A. Krüger, N. Schwarz, R. Rakotozandrindrainy, J. Rakotondrainiarivelo et al., High detection rate of Rickettsia africae in Amblyomma variegatum but low prevalence of anti-rickettsial antibodies in healthy pregnant women in Madagascar, Ticks and Tick-borne Diseases, vol.7, issue.1, pp.60-65, 2016.
DOI : 10.1016/j.ttbdis.2015.08.005

J. Ehlers, J. Ganzhorn, C. Silaghi, A. Krüger, D. Pothmann et al., Tick (Amblyomma chabaudi) infestation of endemic tortoises in southwest Madagascar and investigation of tick-borne pathogens, Ticks and Tick-borne Diseases, vol.7, issue.2, pp.378-83, 2016.
DOI : 10.1016/j.ttbdis.2015.12.011

A. Azad, S. Radulovic, J. Higgins, B. Noden, and J. Troyer, Flea-borne Rickettsioses: Ecologic Considerations, Emerging Infectious Diseases, vol.3, issue.3, pp.319-346, 1997.
DOI : 10.3201/eid0303.970308

A. Azad, Epidemiology of Murine Typhus, Annual Review of Entomology, vol.35, issue.1, pp.553-69, 1990.
DOI : 10.1146/annurev.en.35.010190.003005

H. Dupont, P. Brouqui, B. Faugere, and D. Raoult, Prevalence of Antibodies to Coxiella burnetii, Rickettsia conorii, and Rickettsia typhi in Seven African Countries, Clinical Infectious Diseases, vol.21, issue.5, pp.1126-1159, 1995.
DOI : 10.1093/clinids/21.5.1126

J. Ratovonjato, J. Randriambelosoa, and V. Robert, Tunga penetrans (Insecta, Siphonaptera, Tungidae) à Madagascar: une nuisance négligée, Rev Med Vet, vol.159, pp.551-557, 2008.

P. Graf, J. Chretien, L. Ung, J. Gaydos, and A. Richards, Prevalence of Seropositivity to Spotted Fever Group Rickettsiae and Anaplasma phagocytophilum in a Large, Demographically Diverse US Sample, Clinical Infectious Diseases, vol.46, issue.1, pp.70-71, 2008.
DOI : 10.1086/524018

J. Duchemin, Biogéographie des puces de Madagascar, Thèse de doctorat, 2003.

A. Cornel and F. Collins, PCR of the Ribosomal DNA Intergenic Spacer Regions as a Method for Identifying Mosquitoes in the Anopheles gambiae Complex, Methods Mol Biol, vol.50, pp.321-353, 1996.
DOI : 10.1385/0-89603-323-6:321

C. Socolovschi, O. Mediannikov, C. Sokhna, A. Tall, G. Diatta et al., ???associated Uneruptive Fever, Senegal, Emerging Infectious Diseases, vol.16, issue.7, pp.1140-1142, 2010.
DOI : 10.3201/eid1607.100070

K. Henry, J. Jiang, P. Rozmajzl, A. Azad, K. Macaluso et al., Development of quantitative real-time PCR assays to detect Rickettsia typhi and Rickettsia felis, the causative agents of murine typhus and flea-borne spotted fever, Molecular and Cellular Probes, vol.21, issue.1, pp.17-23, 2007.
DOI : 10.1016/j.mcp.2006.06.002

V. Roux and D. Raoult, Phylogenetic analysis of members of the genus Rickettsia using the gene encoding the outer-membrane protein rOmpB (ompB), INTERNATIONAL JOURNAL OF SYSTEMATIC AND EVOLUTIONARY MICROBIOLOGY, vol.50, issue.4, pp.1449-55, 2000.
DOI : 10.1099/00207713-50-4-1449

J. Jiang, A. Maina, D. Knobel, S. Cleaveland, A. Laudisoit et al., Rickettsia Asemboensis in Fleas from Human Habitats, Asembo, Kenya, Vector-Borne and Zoonotic Diseases, vol.13, issue.8, pp.550-558, 2013.
DOI : 10.1089/vbz.2012.1123

K. Tamura, G. Stecher, D. Peterson, A. Filipski, and S. Kumar, MEGA6: Molecular Evolutionary Genetics Analysis Version 6.0, Molecular Biology and Evolution, vol.30, issue.12, pp.2725-2734, 2013.
DOI : 10.1093/molbev/mst197

R. Team, R: A language and environment for statistical computing, Vienna: R Fund Stat Comput, 2014.

V. Andrianaivoarimanana, K. Kreppel, N. Elissa, J. Duplantier, E. Carniel et al., Understanding the Persistence of Plague Foci in Madagascar, PLoS Neglected Tropical Diseases, vol.5, issue.11, pp.1-8, 2013.
DOI : 10.1371/journal.pntd.0002382.t001

URL : https://hal.archives-ouvertes.fr/pasteur-00946701

A. Richards, D. Soeatmadji, M. Widodo, T. Sardjono, B. Yanuwiadi et al., Seroepidemiologic Evidence for Murine and Scrub Typhus in Malang, Indonesia, The American Journal of Tropical Medicine and Hygiene, vol.57, issue.1, pp.91-96, 1997.
DOI : 10.4269/ajtmh.1997.57.91

A. Maina, D. Knobel, J. Jiang, J. Halliday, D. Feikin et al., Infection in Febrile Patients, Western Kenya, 2007???2010, Emerging Infectious Diseases, vol.18, issue.2, pp.328-359, 2007.
DOI : 10.3201/eid1802.111372

R. Ormsbee, M. Peacock, R. Philip, E. Casper, J. Plorde et al., Antigenic relationship between the typhus and spotted fever group of Rickettsiae, Am J Epidemiol, vol.108, pp.53-62, 1978.

K. Hechemy, D. Raoult, J. Fox, Y. Han, L. Elliott et al., Cross-reaction of immune sera from patients with rickettsial diseases, Journal of Medical Microbiology, vol.29, issue.3, pp.199-202, 1989.
DOI : 10.1099/00222615-29-3-199

A. Znazen, J. Rolain, N. Hammami, A. Hammami, M. Jemaa et al., Infection, Tunisia, Emerging Infectious Diseases, vol.12, issue.1, pp.138-178, 2006.
DOI : 10.3201/eid1201.050876

URL : https://hal.archives-ouvertes.fr/pasteur-01374982

A. Maina, C. Fogarty, L. Krueger, K. Macaluso, A. Odhiambo et al., Rickettsial Infections among Ctenocephalides felis and Host Animals during a Flea-Borne Rickettsioses Outbreak in Orange County, California, PLOS ONE, vol.12, issue.1, p.160604, 2016.
DOI : 10.1371/journal.pone.0160604.s002

K. Mokrani, P. Fournier, M. Dalichaouche, S. Tebbal, A. Aouati et al., Reemerging Threat of Epidemic Typhus in Algeria, Journal of Clinical Microbiology, vol.42, issue.8, pp.3898-900, 2004.
DOI : 10.1128/JCM.42.8.3898-3900.2004

P. Perine, B. Chandler, D. Krause, P. Mccardle, S. Awoke et al., A Clinico-Epidemiological Study of Epidemic Typhus in Africa, Clinical Infectious Diseases, vol.14, issue.5, pp.1149-58, 1992.
DOI : 10.1093/clinids/14.5.1149

J. Berrelha, S. Briolant, F. Muller, J. Rolain, M. J. Pagés et al., Rickettsia felis and Rickettsia massiliae in Ivory Coast, Africa, Clinical Microbiology and Infection, vol.15, pp.251-253, 2009.
DOI : 10.1111/j.1469-0691.2008.02273.x

P. Parola, Rickettsia felis: from a rare disease in the USA to a common cause of fever in sub-Saharan Africa, Clinical Microbiology and Infection, vol.17, issue.7, pp.996-1000, 2011.
DOI : 10.1111/j.1469-0691.2011.03516.x

J. Crump, A. Morrissey, W. Nicholson, R. Massung, R. Stoddard et al., Etiology of Severe Non-malaria Febrile Illness in Northern Tanzania: A Prospective Cohort Study, PLoS Neglected Tropical Diseases, vol.55, issue.7, p.2324, 2013.
DOI : 10.1371/journal.pntd.0002324.s001

A. Richards, J. Jiang, S. Omulo, R. Dare, K. Abdirahman et al., Kenya, Emerging Infectious Diseases, vol.16, issue.7, pp.1081-1087, 2010.
DOI : 10.3201/eid1607.091885

K. Reif and K. Macaluso, Ecology of <I>Rickettsia felis</I>: A Review, Journal of Medical Entomology, vol.46, issue.4, pp.723-759, 2009.
DOI : 10.1603/033.046.0402

M. Eremeeva, W. Warashina, M. Sturgeon, A. Buchholz, G. Olmsted et al., ), Oahu, Hawaii, Emerging Infectious Diseases, vol.14, issue.10, pp.1613-1618, 2008.
DOI : 10.3201/eid1410.080571

J. Jiang, D. Soeatmadji, K. Henry, S. Ratiwayanto, M. Bangs et al., , Java, Indonesia, Emerging Infectious Diseases, vol.12, issue.8, pp.1281-1284, 2006.
DOI : 10.3201/eid1208.060327

C. Christou, A. Psaroulaki, M. Antoniou, P. Toumazos, I. Ioannou et al., Rickettsia typhi and Rickettsia felis in Xenopsylla cheopis and Leptopsylla segnis Parasitizing Rats in Cyprus, The American Journal of Tropical Medicine and Hygiene, vol.83, issue.6, pp.1301-1305, 2010.
DOI : 10.4269/ajtmh.2010.10-0118

C. Sackal, A. Laudisoit, M. Kosoy, R. Massung, M. Eremeeva et al., in Fleas, Democratic Republic of Congo, Emerging Infectious Diseases, vol.14, issue.12, pp.1972-1976, 2008.
DOI : 10.3201/eid1412.080610

A. Maina, A. Luce-fedrow, S. Omulo, J. Hang, T. Chan et al., Isolation and characterization of a new Rickettsia species (Rickettsia asembonensis sp. nov) obtained from cat fleas (Ctenocephalides felis), Int J Syst Evol Microbiol, vol.66, pp.4512-4519, 2016.

O. Mediannikov, C. Socolovschi, S. Edouard, F. Fenollar, N. Mouffok et al., Infection and Malaria, Africa, Emerging Infectious Diseases, vol.19, issue.11, pp.1775-83, 2013.
DOI : 10.3201/eid1911.130361

T. Chareonviriyaphap, W. Leepitakrat, K. Lerdthusnee, C. Chao, and W. Ching, rodents from Thailand, Journal of Vector Ecology, vol.30, issue.1, pp.182-191, 2014.
DOI : 10.1093/ajcp/91.6.720

A. Richards, E. Rahardjo, A. Rusjdi, D. Kelly, G. Dasch et al., Evidence of Rickettsia typhi and the potential for murine typhus in Jayapura, Irian Jaya, Indonesia., The American Journal of Tropical Medicine and Hygiene, vol.66, issue.4, pp.431-435, 2002.
DOI : 10.4269/ajtmh.2002.66.431