J. Beaucournu and D. Fontenille, Contribution à un catalogue des puces de Madagascar (Insecta, Siphonaptera). Archives de l, Institut Pasteur de Madagascar, vol.59, pp.57-98, 1993.

J. Beaucournu, A. R. Randrenjarison, and S. M. Goodman, Puces (Insecta: Siphonaptera) d'Ambohitantely, Madagascar: Spécificité et phénologie. Malagasy Nature, vol.9, pp.39-48, 2015.

E. R. Brygoo, Epidemiologie de la peste à Madagascar. Archives de l, Institut Pasteur de Madagascar, vol.35, pp.9-147, 1966.

T. B. Cheetham, Male genitalia and the phylogeny of the Pulicoidea (Siphonaptera), vol.244, p.pp, 1988.

D. Darriba, G. L. Taboada, R. Doallo, and D. Posada, 2012) jModelTest 2: more models, new heuristics and parallel computing, Nature Methods, vol.9

J. Duchemin, , vol.254, p.pp, 2003.

J. Duchemin, Two new fleas (Siphonaptera: Ceratophyllidae: Leptopsyllinae) of Madagascar: Tsaractenus rodhaini n.sp. and Paractenopsyllus (Consobrinopsyllus n.subgen.) goodmani n.sp. Parasite, vol.10, pp.351-358, 2003.

J. Duchemin, Leptopsyllines from Madagascar (Insecta: Siphonaptera: Ceratophyllidae): description of five new species of Paractenopsyllus Wagner, Parasite, vol.11, pp.59-70, 1938.

J. Duchemin and J. Ratovonjato, Description de trois nouvelles espèces de Paractenopsyllus (Siphonaptera : Leptopsyllinae) de Madagascar. Parasites, vol.11, pp.253-260, 2004.

G. M. Dunnet, D. K. Mardon, P. B. Naumann, J. F. Lawrence, E. S. Nielsen et al., The Insects of Australia: a Textbook for Students and Research Workers, pp.125-140, 1991.

J. Duplantier, J. Duchemin, S. Chanteau, and E. Carniel, From the recent lessons of the Malagasy foci towards a global understanding of the factors involved in plague reemergence, Veterinary Research, vol.36, pp.437-453, 2005.
URL : https://hal.archives-ouvertes.fr/hal-00902976

J. Duplantier, J. B. Duchemin, M. Ratsitorahina, L. Rahalison, and S. Chanteau, Résurgence de la peste dans le district d'Ikongo à Madagascar en 1998. 2. Réservoirs et vecteurs impliqués, vol.94, pp.119-122, 2001.

D. A. Eads, D. E. Biggins, P. F. Doherty, K. L. Gage, K. P. Huyvaert et al., Using occupancy models to investigate the prevalence of ectoparasitic vectors on hosts: An example with fleas on prairie dogs, International Journal for Parasitology: Parasites and Wildlife, vol.2, pp.246-256, 2013.

R. C. Edgar, MUSCLE: multiple sequence alignment with high accuracy and high throughput, Nucleic Acids Research, vol.32, pp.1792-1797, 2004.

G. Girard, Le comportement de la puce Synopsyllus fonquerniei et son rôle dans la transmission de la peste, vol.35, pp.177-181, 1942.

S. M. Goodman, J. U. Ganzhorn, and D. Rakotondravony, Introduction to the mammals, The natural history of Madagascar, pp.1159-1186, 2003.

S. M. Goodman, A. H. Randrenjarison, V. Soarimalala, and J. Beaucournu, The fleas of endemic and introduced small mammals in Central Highland forests of Madagascar: Faunistics, species diversity, and absence of host specificity, Journal of Medical Entomology, vol.52, pp.1135-1143, 2015.


S. M. Goodman, A. P. Raselimanana, and L. Wilmé, Centre d'Information et de Documentation Scientifique et Technique, Anjozorobe-Angavo, vol.24, p.pp, 2007.

S. Guindon and O. Gascuel, A simple, fast, and accurate algorithm to estimate large phylogenies by maximum likelihood, Systematic Biology, vol.52, pp.696-704, 2003.

M. W. Hastriter, Description of two new species of bat fleas of the genus Araeopsylla (Siphonaptera) from Kenya and Madagascar with notes on miscellaneous bat fleas, ZooKeys, vol.572, pp.7-21, 2016.

M. W. Hastriter and C. Dick, A description of the flea species Paractenopsyllus madagascarensis n. sp. and the female sex of Paractenopsyllus raxworthyi Duchemin & Ratovonjato, 2004 (Siphonaptera, Leptopsyllidae) from Madagascar with a key to the species of Paractenopsyllus, ZooKeys, vol.13, pp.43-55, 2009.

S. Kambhampati and P. T. Smith, PCR primers for the amplification of four insect mitochondrial gene fragments, Insect Molecular Biology, vol.4, pp.233-236, 1995.

B. Krasnov, I. Khokhlova, and G. Shenbrot, The effect of host density on ectoparasite distribution: An example of a rodent parasitized by fleas, Ecology, vol.83, pp.164-175, 2002.


A. Laudisoit, V. Prié, and J. Beaucournu, Une nouvelle espèce de Lagaropsylla Jordan & Rothschild, 1921 de Madagascar (Insecta, Siphonaptera, Ischnopsyllidae), Zoosystema, vol.34, pp.737-744, 2012.

R. E. Lewis and J. H. Lewis, Notes on the geographical distribution and host preferences in the order Siphonaptera, Journal of Medical Entomology, vol.22, pp.134-152, 1985.

A. Luchetti, B. Mantovani, S. Pampiglione, and M. Trentini, Molecular characterization of Tunga trimamillata and Tunga penetrans (Insecta, Siphonaptera, Tungidae): taxonomy and genetic variability, Parasite, vol.12, pp.123-129, 2005.

R. Lumaret, Faune de Madagascar: XV: Insectes Siphonaptères. Institut de Recherche Scientifique Madagascar, vol.107, p.pp, 1962.

F. Ronquist, J. Huelsenbeck, and M. Teslenko, Draft MrBayes version 3.2 manual: tutorials and model summaries, vol.172, 2011.

F. Ronquist, M. Teslenko, P. Van-der-mark, D. L. Ayres, A. Darling et al., MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space, Systematic Biology, vol.61, pp.539-542, 2012.

. St, J. R. Juliana, I. S. Khokhlova, N. Wielebnowski, B. P. Kotler et al., Ectoparasitism and stress hormones: strategy of host exploitation, common host-parasite history and energetics matter, Journal of Animal Ecology, vol.83, pp.1113-1123, 2014.

K. Tamura, D. Peterson, N. Peterson, G. Stecher, M. Nei et al., MEGA5: molecular evolutionary genetics analysis using maximum likelihood, evolutionary distance, and maximum parsimony methods, Molecular Biology and Evolution, vol.28, pp.2731-2739, 2011.

J. Wagner and E. Roubaud, Sur un Pulicide nouveau, parasite des rats domestiques à Madagascar, vol.25, pp.327-332, 1932.

J. Wagner and E. Roubaud, Sur un Pulicide nouveau, parasite des rongeurs et insectivores de Madagascar, vol.25, pp.962-964, 1932.

M. F. Whiting, Mecoptera is paraphyletic: multiple genes and phylogeny of Mecoptera and Siphonaptera, Zoologica Scripta, vol.31, pp.93-104, 2002.

M. F. Whiting, A. S. Whiting, M. W. Hastriter, and K. Dittmar, A molecular phylogeny of fleas (Insecta: Siphonaptera): origins and host associations, World Health Statistics, vol.24, pp.677-707, 2008.